Sūtra 7·3·86
पुगन्तलघूपधस्य च
pugantalaghūpadhasya ca
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Guṇa is substituted before a sârvadhâtuka and an ârdbadhâtuka affix, for the ik vowels of the Causative stems which take the augment p (VII.3.36), and for the short penultimate vowel of a root which ends in a single consonant.
Padapāṭha pukantalaghuupadhasya caपुकन्तलघुउपधस्य च