Sūtra 4·1·135
चतुष्पाद्भ्यो ढञ्
catuṣpādbhyo ḍhañ
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The affix ‘ḍhañ’ comes in the sense of a descendant, after the bases denoting quadrupeds.
Padapāṭha catuṣpādbhyaḥ ḍhañचतुष्पाद्भ्यः ढञ्
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